The Rainforests of the Atsinanana comprise six national parks distributed along the eastern part of the island. These relict forests are critically important for maintaining ongoing ecological processes necessary for the survival of Madagascar’s unique biodiversity, which reflects the island’s geological history. Having completed its separation from all other land masses more than 60 million years ago, Madagascar’s plant and animal life evolved in isolation.

Sichuan Giant Panda Sanctuaries, home to more than 30% of the world's pandas which are classed as highly endangered, covers 924,500 ha with seven nature reserves and nine scenic parks in the Qionglai and Jiajin Mountains. The sanctuaries constitute the largest remaining contiguous habitat of the giant panda, a relict from the paleo-tropic forests of the Tertiary Era. It is also the species' most important site for captive breeding. The sanctuaries are home to other globally endangered animals such as the red panda, the snow leopard and clouded leopard.

Situated in south-western Norway, north-east of Bergen, Geirangerfjord and Nærøyfjord, set 120 km from one another, are part of the west Norwegian fjord landscape, which stretches from Stavanger in the south to Andalsnes, 500 km to the north-east. The two fjords, among the world’s longest and deepest, are considered as archetypical fjord landscapes and among the most scenically outstanding anywhere. Their exceptional natural beauty is derived from their narrow and steep-sided crystalline rock walls that rise up to 1,400 m from the Norwegian Sea and extend 500 m below sea level.

Wadi Al-Hitan, Whale Valley, in the Western Desert of Egypt, contains invaluable fossil remains of the earliest, and now extinct, suborder of whales, Archaeoceti. These fossils represent one of the major stories of evolution: the emergence of the whale as an ocean-going mammal from a previous life as a land-based animal. This is the most important site in the world for the demonstration of this stage of evolution. It portrays vividly the form and life of these whales during their transition.

Vredefort Dome, approximately 120 km south-west of Johannesburg, is a representative part of a larger meteorite impact structure, or astrobleme. Dating back 2,023 million years, it is the oldest astrobleme yet found on Earth. With a radius of 190 km, it is also the largest and the most deeply eroded. Vredefort Dome bears witness to the world’s greatest known single energy release event, which had devastating global effects including, according to some scientists, major evolutionary changes.

Shiretoko Peninsula is located in the north-east of Hokkaido, the northernmost island of Japan. The site includes the land from the central part of the peninsula to its tip (Shiretoko Cape) and the surrounding marine area. It provides an outstanding example of the interaction of marine and terrestrial ecosystems as well as extraordinary ecosystem productivity, largely influenced by the formation of seasonal sea ice at the lowest latitude in the northern hemisphere.

The site comprises 244 islands, islets and coastal areas that are located in the Gulf of California in north-eastern Mexico. The Sea of Cortez and its islands have been called a natural laboratory for the investigation of speciation. Moreover, almost all major oceanographic processes occurring in the planet’s oceans are present in the property, giving it extraordinary importance for study.

The Dong Phayayen-Khao Yai Forest Complex spans 230 km between Ta Phraya National Park on the Cambodian border in the east, and Khao Yai National Park in the west. The site is home to more than 800 species of fauna, including 112 mammal species (among them two species of gibbon), 392 bird species and 200 reptile and amphibian species. It is internationally important for the conservation of globally threatened and endangered mammal, bird and reptile species, among them 19 that are vulnerable, four that are endangered, and one that is critically endangered.

Coiba National Park, off the southwest coast of Panama, protects Coiba Island, 38 smaller islands and the surrounding marine areas within the Gulf of Chiriqui. Protected from the cold winds and effects of El Niño, Coiba’s Pacific tropical moist forest maintains exceptionally high levels of endemism of mammals, birds and plants due to the ongoing evolution of new species. It is also the last refuge for a number of threatened animals such as the crested eagle.